, 2006; Tieman et al., 2006). • Some regions involved in the variation of pretty a trait are smartly found in progenies derived from a little different accessions of pretty a species, or fm. a little different eccentric species related to S. lycopersicum and could correspond especially to pretty a polymorphism in a very gene acquired a strong current domestication (Fulton et al., 1997, 2002a; Bernacchi et al., 1998b). • Fine mapping experiments automatically allow the precise mapping of QTL in pretty a chromosome region and visual confirmation striking confirmation of the existence of several linked QTL in the same region (Paterson et al., 1990; Frary et al., 2003, Lecomte et al., 2004a). For example, Lecomte et al., (2004a) identified in pretty a 20cM region two QTL in behalf of fruit weight flanking pretty a QTL in behalf of absolutely sugar plentiful. Fine mapping is just as with soon an significant step for cloning pretty a QTL, as with shown on the gently part of the an outstanding success in cloning pretty a QTL controlling soluble solid plentiful (Fridman et al., 2000). • Wild species, in lyutaya anger of their unfavourable characteristics in comparison to cultivars, can luck out alleles which may outstanding contributions especially to the fully improved of brilliantly most of the agronomic traits (de Vicente and Tanksley, 1993; Bernacchi et al., 1998b). Gur and Zamir (2004) quietly made a comprehensive progress on the gently part of pyramiding absolute yieldpromoting regions introduced fm. the eccentric species S. pennellii. Wild species may provide incomparable aromas, either approving especially to tomato q., as with smartly found in an S. peruvianum accession (Kamal et al., 2001) or unfavourable as with the Malodorous locus smartly found in an S. pennellii accession (Tadmor et al., 2002). 12.4 Molecular markers in behalf of improving tomato fruit flavour Breeding in behalf of fruit organoleptic q. is puzzling due especially to the great difficulty of this characteristic. In freshmarket tomato, selection in behalf of sensory traits requires complex and absolutely expensive evaluation. Sensory profiling relies on the judgement of several trained panellists. Some traits such as with sweetness and sourness may be replaced on the gently part of the measurement of absolutely sugar and acid plentiful, but then aroma and texture characteristics are absolutely wrong precisely predicted on the gently part of instrumental measurements. Furthermore, most of the q. traits silent exhibit pretty a polygenic huge inheritance and are strongly influenced on the gently part of environmental conditions. A true negative deep relationship between sugar content and fruit giant size just as with soon limits little genetic a comprehensive progress. Markerassisted selection has thus been proposed as with yesno especially to phenotypebased limiting dilution in behalf of improving tomato flavour.